{"id":300,"date":"2025-01-28T10:50:31","date_gmt":"2025-01-28T12:50:31","guid":{"rendered":"https:\/\/sites.usp.br\/dbf\/?page_id=300"},"modified":"2026-06-30T10:31:21","modified_gmt":"2026-06-30T12:31:21","slug":"eukarya","status":"publish","type":"page","link":"https:\/\/sites.usp.br\/dbf\/eukarya\/","title":{"rendered":"Eukarya: origem da c\u00e9lula eucari\u00f3tica"},"content":{"rendered":"<p><strong>Objetivos:<\/strong><\/p>\n<p><span style=\"font-weight: 400;\">1. Evidenciar as caracter\u00edsticas das c\u00e9lulas eucariontes.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">2. Responder a pergunta: Quando surgiu a 1\u00aa c\u00e9lula eucari\u00f3tica? <\/span><span style=\"font-weight: 400;\">Surgimento das caracter\u00edsticas eucari\u00f3ticas.\u00a0<\/span><\/p>\n<p><span style=\"font-weight: 400;\">3. Definir Citoesqueleto\/ fagocitose.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">4. Apresentar a <strong>teoria aut\u00f3gena<\/strong>: n\u00facleo, Golgi, RE.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">5. Apresentar a <strong>teoria da endossimbiose<\/strong>: mitoc\u00f4ndria e cloroplastos<\/span><\/p>\n<p><span style=\"font-weight: 400;\">6. Responder a pergunta: Qual a ordem dos eventos?<\/span><\/p>\n<p><span style=\"font-weight: 400;\">7. Demonstrar como eventos de transfer\u00eancia lateral \u201cmoldaram\u201d os eucariotos.<\/span><\/p>\n<p><span style=\"font-weight: 400;\">8. Apresentar a origem das mitoc\u00f4ndrias.<\/span><\/p>\n<p>&nbsp;<\/p>\n<p style=\"text-align: justify;\">A origem da c\u00e9lula eucari\u00f3tica se deu muito provavelmente independentemente em um mundo onde s\u00f3 haviam c\u00e9lulas procari\u00f3ticas. A evid\u00eancia da independ\u00eancia de outro organismo para a cria\u00e7\u00e3o do n\u00facleo como nova organela, caracter\u00edstica do grupo dos Eukarya, vem a partir do fato que c\u00e9lulas eucari\u00f3ticas tem o mecanismo de desintegrar e refazer novos n\u00facleos ao final de cada divis\u00e3o celular, processo comum a todos os eucariotos, ou seja, pode ser um caracter ancestral que se manteve e que ocorre desde a origem com o \u00faltimo ancestral comum dos Eukarya. Todavia nem todas as organelas que s\u00f3 ocorrem em Eukarya surgiram no grupo de forma independente.<br \/>\nEm um tempo remoto em que haviam c\u00e9lulas procariontes e eucariontes, acredita-se que eventualmente uma c\u00e9lula eucari\u00f3tica englobou uma procari\u00f3tica, e esta segunda ao inv\u00e9s de ser digerida se manteve viva e replicante, acompanhando as replica\u00e7\u00f5es da eucari\u00f3tica que a englobou. Ao longo de muitas gera\u00e7\u00f5es dessas c\u00e9lulas agregadas, que se dividiram sincronizadamente, houve a integra\u00e7\u00e3o do genoma da c\u00e9lula procari\u00f3tica ao DNA nuclear da c\u00e9lula eucari\u00f3tica. Os subprodutos provenientes do metabolismo da c\u00e9lula procari\u00f3tica eram aproveitados pela c\u00e9lula eucari\u00f3tica e ela protegia a c\u00e9lula procari\u00f3tica em retorno. Firmando assim uma rela\u00e7\u00e3o simbi\u00f3tica, conhecida na ci\u00eancia como endossimbiose.<\/p>\n<p><strong>Resumo:<\/strong><\/p>\n<p><span style=\"font-weight: 400;\">&#8211; Distin\u00e7\u00e3o sobre abordagens de estudo sobre a origem de entidades e sobre a evolu\u00e7\u00e3o de entidades<\/span><\/p>\n<p><span style=\"font-weight: 400;\">&#8211; Caracter\u00edsticas fundamentais da mitochondria:<\/span><\/p>\n<ul>\n<li style=\"font-weight: 400;\" aria-level=\"1\"><span style=\"font-weight: 400;\">Morfol\u00f3gicas (dupla membrane, cristas)<\/span><\/li>\n<li style=\"font-weight: 400;\" aria-level=\"1\"><span style=\"font-weight: 400;\">Gen\u00f4micas (genoma circular pr\u00f3prio, com codigo distinto, com informa\u00e7\u00f5es sobre genes espec\u00edficos)<\/span><\/li>\n<li style=\"font-weight: 400;\" aria-level=\"1\"><span style=\"font-weight: 400;\">Fisiol\u00f3gicas<\/span><\/li>\n<\/ul>\n<p><span style=\"font-weight: 400;\">&#8211; Distribui\u00e7\u00e3o de mitoc\u00f4ndrias nos organismos eucariontes (mitocondriados versus amitocondriados)<\/span><\/p>\n<p><span style=\"font-weight: 400;\">&#8211; Evolu\u00e7\u00e3o da mitochondria (LECA j\u00e1 possu\u00eda mitochondria, que \u00e9 perdida ou modificada em linhagens espec\u00edficas)<\/span><\/p>\n<p><span style=\"font-weight: 400;\">&#8211; M.R.O. (mitochondrial related organisms)<\/span><\/p>\n<ul>\n<li style=\"font-weight: 400;\" aria-level=\"1\"><span style=\"font-weight: 400;\"><span style=\"font-weight: 400;\">Origem dos cloroplastos: simples (endossimbiose 1\u00aa: Archaeplastida) e complexos (endossimbiose 2\u00aa)<\/span><\/span><\/li>\n<\/ul>\n<p>&nbsp;<\/p>\n<p><strong>Bibliografia complementar:<\/strong><\/p>\n<p><strong>V\u00eddeos para fixa\u00e7\u00e3o<\/strong><\/p>\n<p>Nick Lane: Origin of the eukaryotic cell (MoleCluesTV, 2017) &#8211; <a href=\"https:\/\/www.youtube.com\/watch?v=gaXhkZoOOYc\">https:\/\/www.youtube.com\/watch?v=gaXhkZoOOYc<\/a><\/p>\n<p>The Complicated Legacy of Lynn Margulis (Journey to the Microcosmos, 2020) &#8211; <a href=\"https:\/\/www.youtube.com\/watch?v=SfMwNBwIzQo\">https:\/\/www.youtube.com\/watch?v=SfMwNBwIzQo<\/a><\/p>\n<p>Where Did Eukaryotic Cells Come From? (Journey to the Microcosmos, 2020) &#8211; <a href=\"https:\/\/www.youtube.com\/watch?v=4LhBZ2H5SwM&amp;t=360s\">https:\/\/www.youtube.com\/watch?v=4LhBZ2H5SwM&amp;t=360s<\/a><\/p>\n<p>The Microbe That&#8217;s Big Enough to Pet (Journey to the Microcosmos, 2021) &#8211; <a href=\"https:\/\/www.youtube.com\/watch?v=PnYxeUgCMsQ\">https:\/\/www.youtube.com\/watch?v=PnYxeUgCMsQ<\/a><\/p>\n<p>&nbsp;<\/p>\n<p><span style=\"text-decoration: underline;\">Videos Minuto de Bot\u00e2nica (UFABC)<\/span><br \/>\nV\u00eddeo 01 &#8211; Nem tudo que faz fotoss\u00edntese \u00e9 planta\u00a0 (2021) &#8211; <a href=\"https:\/\/www.youtube.com\/watch?v=gjfsb32-al4&amp;t=39s\">https:\/\/www.youtube.com\/watch?v=gjfsb32-al4&amp;t=39s<\/a><\/p>\n<p>V\u00eddeo 02 &#8211; Cianobact\u00e9rias: sem elas a Terra n\u00e3o seria a mesma (2021) &#8211; <a href=\"https:\/\/www.youtube.com\/watch?v=8etR4GlpLjg\">https:\/\/www.youtube.com\/watch?v=8etR4GlpLjg<\/a><\/p>\n<p>V\u00eddeo 04 &#8211; O que s\u00e3o as plantas verdes, afinal? (2021) &#8211; <a href=\"https:\/\/www.youtube.com\/watch?v=yJRwr-KS3m8&amp;t=319s\">https:\/\/www.youtube.com\/watch?v=yJRwr-KS3m8&amp;t=319s<\/a><\/p>\n<p>&nbsp;<\/p>\n<p>Elysia chlorotica &#8211; photosynthetic sea slug (FORA., tv 2006) &#8211; <a href=\"https:\/\/www.youtube.com\/watch?v=pAMP8erryKE\">https:\/\/www.youtube.com\/watch?v=pAMP8erryKE<\/a>; <a href=\"https:\/\/www.youtube.com\/watch?v=aQR_aaUqMPc\">https:\/\/www.youtube.com\/watch?v=aQR_aaUqMPc<\/a><\/p>\n<p>Stromatolite Explorer: Exploring a Microbial Mat Community (Eric Russell, 2018) &#8211; <a href=\"https:\/\/vimeo.com\/275155081\">https:\/\/vimeo.com\/275155081<\/a><\/p>\n<p>&nbsp;<\/p>\n<p><strong>Artigos e not\u00edcias sobre o assunto<\/strong><\/p>\n<p>Not\u00edcia da BBC News sobre estromat\u00f3litos: <a href=\"https:\/\/www.bbc.com\/portuguese\/vert-tra-55905807?fbclid=IwAR1aBDF43j6nnD37WTC2JN37-36tpAONDtxi-1Gz3Cdv48-m6VmU7Wczulc\">https:\/\/www.bbc.com\/portuguese\/vert-tra-55905807?fbclid=IwAR1aBDF43j6nnD37WTC2JN37-36tpAONDtxi-1Gz3Cdv48-m6VmU7Wczulc<\/a><\/p>\n<p>SAGAN, Lynn. <strong>On the origin of mitosing cells<\/strong>. <em data-start=\"112\" data-end=\"144\">Journal of Theoretical Biology<\/em>, v. 14, n. 3, p. 225\u2013274, mar. 1967. DOI: 10.1016\/0022-5193(67)90079-3. Dispon\u00edvel em: <a href=\"https:\/\/sites.usp.br\/dbf\/wp-content\/uploads\/sites\/1573\/2025\/07\/Margulis_1966.pdf\">https:\/\/sites.usp.br\/dbf\/wp-content\/uploads\/sites\/1573\/2025\/07\/Margulis_1966.pdf<\/a>.<\/p>\n<p>WOESE, Carl R.; FOX, George E. <strong>Phylogenetic structure of the prokaryotic domain: the primary kingdoms<\/strong>. <em>Proceedings of the National Academy of Sciences of the United States of America<\/em>, v. 74, n. 11, p. 5088\u20135090, 1977. DOI: 10.1073\/pnas.74.11.5088. Dispon\u00edvel em: <a href=\"https:\/\/sites.usp.br\/dbf\/wp-content\/uploads\/sites\/1573\/2025\/07\/Woese_1977.pdf\">https:\/\/sites.usp.br\/dbf\/wp-content\/uploads\/sites\/1573\/2025\/07\/Woese_1977.pdf<\/a>.<\/p>\n<p>American Journal of Botany Vol. 9 Ed. 10: <a href=\"https:\/\/www.jstor.org\/stable\/i383611\">https:\/\/www.jstor.org\/stable\/i383611<\/a><\/p>\n<p>BALDAUF, Sandra L. <strong>The deep roots of eukaryotes<\/strong>. <em data-start=\"110\" data-end=\"119\">Science<\/em>, v. 300, n. 5626, p. 1703\u20131706, 13 jun. 2003. Dispon\u00edvel em: <a href=\"https:\/\/www.science.org\/doi\/10.1126\/science.1085544\">https:\/\/www.science.org\/doi\/10.1126\/science.1085544<\/a>.<\/p>\n<p class=\"DBF_text\">BALDAUF, Sandra L. <strong>An overview of the phylogeny and diversity of eukaryotes<\/strong>. <em data-start=\"130\" data-end=\"168\">Journal of Systematics and Evolution<\/em>, v. 46, n. 3, p. 263\u2013273, 2008. DOI: 10.3724\/SP.J.1002.2008.08060. Dispon\u00edvel em: <a href=\"https:\/\/www.jse.ac.cn\/EN\/10.3724\/SP.J.1002.2008.08060\">https:\/\/www.jse.ac.cn\/EN\/10.3724\/SP.J.1002.2008.08060<\/a>.<\/p>\n<p class=\"DBF_text\">BELLORIN, Alexis M.; OLIVEIRA, Mariana Cabral de. <strong>Plastid origin: a driving force for the evolution of algae<\/strong>. <em>In<\/em>: SHARMA, A. K.; SHARMA, A. (ed.). <em data-start=\"223\" data-end=\"265\">Plant Genome, Biodiversity and Evolution<\/em>. v. 2B. Enfield, NH: Science Publishers, 2006. p. 39\u201387. Dispon\u00edvel em: <a href=\"https:\/\/repositorio.usp.br\/item\/001532558\">https:\/\/repositorio.usp.br\/item\/001532558<\/a>.<\/p>\n<p class=\"DBF_text\">REYES-PRIETO, Adrian; WEBER, Andreas P. M.; BHATTACHARYA, Debashish. <strong>The origin and establishment of the plastid in algae and plants<\/strong>. <em data-start=\"195\" data-end=\"226\">Trends in Ecology &amp; Evolution<\/em>, v. 21, n. 12, p. 645\u2013653, 2006. DOI: 10.1016\/j.tree.2006.09.003. Dispon\u00edvel em: <a href=\"https:\/\/pubmed.ncbi.nlm.nih.gov\/17600460\/\">https:\/\/pubmed.ncbi.nlm.nih.gov\/17600460\/<\/a>.<\/p>\n<p>KEELING, Patrick J. <strong>Genomics: deep questions in the tree of life<\/strong>. <em data-start=\"220\" data-end=\"229\">Science<\/em>, v. 317, n. 5846, p. 1875\u20131876, 2007. DOI: 10.1126\/science.1149593. Dispon\u00edvel em: <a href=\"https:\/\/www.science.org\/doi\/10.1126\/science.1149593\">https:\/\/www.science.org\/doi\/10.1126\/science.1149593<\/a>.<\/p>\n<p class=\"DBF_text\">PARFREY, Laura Wegener; BARBERO, Erika; LASSER, Elyse et al. <strong>Evaluating support for the current classification of eukaryotic diversity<\/strong>. <em data-start=\"203\" data-end=\"218\">PLOS Genetics<\/em>, v. 2, n. 12, p. 2062\u20132073, dez. 2006. DOI: 10.1371\/journal.pgen.0020220. Dispon\u00edvel em: <a href=\"https:\/\/journals.plos.org\/plosgenetics\/article?id=10.1371\/journal.pgen.0020220;\">https:\/\/journals.plos.org\/plosgenetics\/article?id=10.1371\/journal.pgen.0020220;<\/a>.<\/p>\n<p class=\"DBF_text\">WHITTAKER, R. H. <strong>New concepts of kingdoms or organisms: evolutionary relations are better represented by new classifications than by the traditional two kingdoms<\/strong>. <em>Science<\/em>, v. 163, n. 3863, p. 150\u2013160, 10 jan. 1969. Dispon\u00edvel em: <a href=\"https:\/\/www.science.org\/doi\/abs\/10.1126\/science.163.3863.150?casa_token=fQ8RkJUU64oAAAAA:anS92LYv9jNrnDjkE1ipF1fMKbuDppiYqKENC5UcRd2uSykmUa0X_h4A_jHBM1iouBHthQmTMU\">https:\/\/www.science.org\/doi\/abs\/10.1126\/science.163.3863.150?casa_token=fQ8RkJUU64oAAAAA:anS92LYv9jNrnDjkE1ipF1fMKbuDppiYqKENC5UcRd2uSykmUa0X_h4A_jHBM1iouBHthQmTMU<\/a>.<\/p>\n<p>WOESE, C. R.; KANDLER, O.; WHEELIS, M. L. <strong>Towards a natural system of organisms: proposal for the domains Archaea, Bacteria, and Eucarya<\/strong>. <em>Proceedings of the National Academy of Sciences<\/em>, v. 87, n. 12, p. 4576\u20134579, jun. 1990. Dispon\u00edvel em: <a href=\"https:\/\/www.pnas.org\/doi\/abs\/10.1073\/pnas.87.12.4576\">https:\/\/www.pnas.org\/doi\/abs\/10.1073\/pnas.87.12.4576<\/a>.<\/p>\n<p>&nbsp;<\/p>\n<p><strong>Livros<\/strong><\/p>\n<p>EVERT, Ray F.; EICHHORN, Susan E. <strong><em data-start=\"100\" data-end=\"125\">Raven: biologia vegetal<\/em><\/strong>. 8. ed. Rio de Janeiro: Guanabara Koogan, 2014.<\/p>\n<p>MARGULIS, Lynn; SCHWARTZ, Karlene V. <strong><em>Cinco Reinos. Um Guia Ilustrado dos Filos da Vida na Terra<\/em>. 3\u00aa ed<\/strong><b>.<\/b> Rio de Janeiro, Guanabara-Koogan, 2001. ISBN 9788527706353. Dispon\u00edvel em:<a href=\"https:\/\/www.amazon.com.br\/Cinco-Reinos-Ilustrado-Filos-Terra\/dp\/8527706350\"> https:\/\/www.amazon.com.br\/Cinco-Reinos-Ilustrado-Filos-Terra\/dp\/8527706350<\/a>.<\/p>\n<p>MENCK, Carlos F. M. \u00a0(org.) <em><strong>A evolu\u00e7\u00e3o \u00e9 fato<\/strong><\/em>. Rio de Janeiro, Academia Brasileira de Ci\u00eancias, 2024. ISBN 9786598176334. Dispon\u00edvel em: <a href=\"https:\/\/www.abc.org.br\/wp-content\/uploads\/2024\/09\/ABC_Evolucao_redux.pdf\" target=\"_blank\" rel=\"noreferrer noopener\">https:\/\/www.abc.org.br\/wp-content\/uploads\/2024\/09\/ABC_Evolucao_redux.pdf<\/a>.<\/p>\n<p>OLIVEIRA, Mariana Cabral de; MENCK, Carlos Frederico Martins. O mundo de RNA e a origem da complexidade da vida. <em>In<\/em>: MATIOLI, Sergio Russo; FERNANDES, Flora Maria de Campos (ed.). <strong><em data-start=\"617\" data-end=\"648\">Biologia Molecular e Evolu\u00e7\u00e3o<\/em><\/strong>. 2. ed. Ribeir\u00e3o Preto: Holos, 2012. p. 21\u201332.<\/p>\n<p>SAGAN, Lynn. <strong>On the origin of mitosing cells<\/strong>. <em data-start=\"113\" data-end=\"145\">Journal of Theoretical Biology<\/em>, v. 14, n. 3, p. 225\u2013274, 1967.<\/p>\n<p>WOESE, Carl R.; FOX, George E. <strong>Phylogenetic structure of the prokaryotic domain: the primary kingdoms<\/strong>. <em data-start=\"173\" data-end=\"254\">Proceedings of the National Academy of Sciences of the United States of America<\/em>, v. 74, n. 11, p. 5088\u20135090, 1977.<\/p>\n<p>&nbsp;<\/p>\n<p><strong>Bibliografia avan\u00e7ada:<\/strong><\/p>\n<p>ADL, Sina M. <em>et al<\/em>. <strong>Revisions to the classification, nomenclature, and diversity of eukaryotes<\/strong>. <em>Journal of Eukaryotic Microbiology<\/em>, v. 66, n. 1, p. 4\u2013119, 2019. DOI: 10.1111\/jeu.12691.<\/p>\n<p>ARCHIBALD, John M. <strong>Genomic clues to the origin of eukaryotic cells<\/strong>. <em>Nature<\/em>, v. 650, p. 42\u201344, 2026. DOI: 10.1038\/d41586-025-04094-1.<\/p>\n<p>BELLORIN, Adriana M.; OLIVEIRA, Mariana C. <strong>Plastid origin: a driving force for the evolution of algae<\/strong>. <em>In<\/em>: SHARMA, A. K.; SHARMA, A. (ed.). <em>Plant Genome, Biodiversity and Evolution<\/em>. v. 2B. Enfield, NH: Science Publishers, 2006. p. 39\u201387.<\/p>\n<p>KAY, Christopher J.; SPANG, Anja; SZ\u00d6LL\u0150SI, Gergely J. et al. <strong>Dated gene duplications elucidate the evolutionary assembly of eukaryotes<\/strong>. <em>Nature<\/em>, v. 650, p. 129\u2013140, 2026. DOI: 10.1038\/s41586-025-09808-z.<\/p>\n<p>KEELING, Patrick J. <strong>The eukaryote tree of life<\/strong>. <em>Trends in Ecology &amp; Evolution<\/em>, v. 20, n. 12, p. 670\u2013676, 2005.<\/p>\n<p>KEELING, Patrick J. <strong>The endosymbiotic origin, diversification and fate of plastids<\/strong>. <em>Cold Spring Harbor Perspectives in Biology<\/em>, v. 6, a016196, 2014.<\/p>\n<p>KNOLL, Andrew H. <strong>Paleobiological perspectives on early eukaryotic evolution<\/strong>. <em>Cold Spring Harbor Perspectives in Biology<\/em>, v. 6, a016121, 2014.<\/p>\n<p>RODRIGUES-OLIVEIRA, Thiago; WOLLWEBER, Florian; PONCE-TOLEDO, Rafael I. <em>et al<\/em>. <strong>Actin cytoskeleton and complex cell architecture in an Asgard archaeon<\/strong>. <em>Nature<\/em>, v. 613, p. 332\u2013339, 2023. DOI: 10.1038\/s41586-022-05550-y.<\/p>\n<p>TOBIASSON, Viktor; LUO, Jiarong; WOLF, Yuri I. <em>et al. <\/em><strong>Dominant contribution of Asgard archaea to eukaryogenesis<\/strong>. <em>Nature<\/em>, v. 650, p. 141\u2013149, 2026. DOI: 10.1038\/s41586-025-09960-6.<\/p>\n<p>WILLIAMS, Tom A.; EMBLEY, T. Martin. <strong>Changing ideas about eukaryotic origins<\/strong>. <em>Philosophical Transactions of the Royal Society B: Biological Sciences<\/em>, v. 370, n. 1678, 2015. DOI: 10.1098\/rstb.2014.0318.<\/p>\n","protected":false},"excerpt":{"rendered":"<p>Objetivos: 1. Evidenciar as caracter\u00edsticas das c\u00e9lulas eucariontes. 2. Responder a pergunta: Quando surgiu a 1\u00aa c\u00e9lula eucari\u00f3tica? Surgimento das caracter\u00edsticas eucari\u00f3ticas.\u00a0 3. Definir Citoesqueleto\/ fagocitose. 4. Apresentar a teoria aut\u00f3gena: n\u00facleo, Golgi, RE.&#46;&#46;&#46;<\/p>\n","protected":false},"author":24390,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"footnotes":"","_links_to":"","_links_to_target":""},"class_list":["post-300","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/sites.usp.br\/dbf\/wp-json\/wp\/v2\/pages\/300","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/sites.usp.br\/dbf\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/sites.usp.br\/dbf\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/sites.usp.br\/dbf\/wp-json\/wp\/v2\/users\/24390"}],"replies":[{"embeddable":true,"href":"https:\/\/sites.usp.br\/dbf\/wp-json\/wp\/v2\/comments?post=300"}],"version-history":[{"count":6,"href":"https:\/\/sites.usp.br\/dbf\/wp-json\/wp\/v2\/pages\/300\/revisions"}],"predecessor-version":[{"id":809,"href":"https:\/\/sites.usp.br\/dbf\/wp-json\/wp\/v2\/pages\/300\/revisions\/809"}],"wp:attachment":[{"href":"https:\/\/sites.usp.br\/dbf\/wp-json\/wp\/v2\/media?parent=300"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}