{"id":868,"date":"2025-02-15T23:49:16","date_gmt":"2025-02-16T02:49:16","guid":{"rendered":"https:\/\/sites.usp.br\/lesca\/?page_id=868"},"modified":"2025-04-08T20:29:41","modified_gmt":"2025-04-08T23:29:41","slug":"comportamento-animal","status":"publish","type":"page","link":"https:\/\/sites.usp.br\/lesca\/comportamento-animal\/","title":{"rendered":"Comportamento animal"},"content":{"rendered":"\t\t<div data-elementor-type=\"wp-page\" data-elementor-id=\"868\" class=\"elementor elementor-868\">\n\t\t\t\t\t\t<section class=\"elementor-section elementor-top-section elementor-element elementor-element-87f42d5 elementor-section-boxed elementor-section-height-default elementor-section-height-default\" data-id=\"87f42d5\" data-element_type=\"section\">\n\t\t\t\t\t\t<div class=\"elementor-container elementor-column-gap-default\">\n\t\t\t\t\t<div class=\"elementor-column elementor-col-50 elementor-top-column elementor-element elementor-element-dbb6551\" data-id=\"dbb6551\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-731488f elementor-widget elementor-widget-text-editor\" data-id=\"731488f\" data-element_type=\"widget\" data-widget_type=\"text-editor.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t\t\t<p>\u00a0 \u00a0<b>Estudo do comportamento animal e o LESCA<\/b><br \/>Neste texto farei um brev\u00edssimo hist\u00f3rico de estudos sobre comportamento animal e ao final tentarei encaixar o LESCA nesse contexto em termos de abordagem e m\u00e9todos.<\/p><p>\u00a0 \u00a0Do s\u00e9culo XVIII at\u00e9 o s\u00e9culo XIX, duas escolas de pensamento discutiram avidamente sobre qual a maneira mais adequada de se estudar o comportamento animal. Fisiologistas como <b>Charles Bell<\/b> (1774-1842), <b>Fran\u00e7ois Magendie<\/b> (1783-1855) e <b>Pierre Flourens<\/b> (1794-1867) foram dos precursores da escola mecanista, um grupo de pensadores que defendia arduamente o empiricismo e a realiza\u00e7\u00e3o de experimentos. Eles davam continuidade a uma linha de pensamento de certa forma iniciada por <b>Ren\u00e9 Descartes<\/b> (1596-1650) que iria influenciar cientistas por d\u00e9cadas, dentre os quais <b>Jacques Loeb<\/b> (1859-1924) e <b>Ivan Pavlov <\/b>(1849-1936). Tais cientistas buscavam encontrar leis do comportamento, preocupando-se principalmente com quest\u00f5es proximais e do desenvolvimento. Essa escola defendia o rigoroso controle das condi\u00e7\u00f5es experimentais e para isso a pesquisa deveria necessariamente ser feita em laborat\u00f3rio. Deles veio o grupo da \u201cPsicologia Experimental\u201d, ao qual tamb\u00e9m pertenciam <b>William James <\/b>(1842-1910), <b>Edward Thorndike<\/b> (1874-1949),<b> John Watson<\/b> (1878-1958) e <b>Burrhus Frederic Skinner <\/b>(1904-1990). Estes ficaram famosos por seus experimentos de condicionamento e aprendizagem como um todo, utilizando vertebrados como modelos.<\/p><div>\u00a0<\/div>\t\t\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t<div class=\"elementor-column elementor-col-50 elementor-top-column elementor-element elementor-element-be837a5\" data-id=\"be837a5\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-4772293 elementor-widget__width-initial elementor-widget elementor-widget-image\" data-id=\"4772293\" data-element_type=\"widget\" data-widget_type=\"image.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t\t\t\t\t\t\t\t\t<img fetchpriority=\"high\" decoding=\"async\" width=\"1000\" height=\"667\" src=\"https:\/\/sites.usp.br\/lesca\/wp-content\/uploads\/sites\/1615\/2025\/04\/opilion.jpg\" class=\"attachment-full size-full wp-image-1405\" alt=\"\" srcset=\"https:\/\/sites.usp.br\/lesca\/wp-content\/uploads\/sites\/1615\/2025\/04\/opilion.jpg 1000w, https:\/\/sites.usp.br\/lesca\/wp-content\/uploads\/sites\/1615\/2025\/04\/opilion-300x200.jpg 300w, https:\/\/sites.usp.br\/lesca\/wp-content\/uploads\/sites\/1615\/2025\/04\/opilion-768x512.jpg 768w, https:\/\/sites.usp.br\/lesca\/wp-content\/uploads\/sites\/1615\/2025\/04\/opilion-400x267.jpg 400w\" sizes=\"100vw\" \/>\t\t\t\t\t\t\t\t\t\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/section>\n\t\t\t\t<section class=\"elementor-section elementor-top-section elementor-element elementor-element-9066067 elementor-section-boxed elementor-section-height-default elementor-section-height-default\" data-id=\"9066067\" data-element_type=\"section\">\n\t\t\t\t\t\t<div class=\"elementor-container elementor-column-gap-default\">\n\t\t\t\t\t<div class=\"elementor-column elementor-col-100 elementor-top-column elementor-element elementor-element-cebc610\" data-id=\"cebc610\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-a2d5edb elementor-widget elementor-widget-text-editor\" data-id=\"a2d5edb\" data-element_type=\"widget\" data-widget_type=\"text-editor.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t\t\t<p>\u00a0 \u00a0Paralelamente a este grupo, desenvolvia-se uma maneira diferente de estudar o comportamento. Eram pessoas que observavam os animais durante suas caminhadas no campo e eventualmente faziam anota\u00e7\u00f5es. Menos preocupados com leis do comportamento que necessariamente valessem para qualquer t\u00e1xon, procuravam muitas vezes exatamente as diferen\u00e7as entre as esp\u00e9cies. Estes eram os chamados naturalistas, dentre os quais podemos destacar o <b>Comte de Buffon<\/b> (1707-1788), <b>Fritz Muller <\/b>(1821-1897), <b>Henry Bates<\/b> (1825-1892) e o pr\u00f3prio <b>Charles Darwin<\/b> (1809-1882), entre outros. Um marco hist\u00f3rico neste grupo veio com <b>Nikolaas Tinbergen<\/b> (1907-1988), <b>Karl von Frisch<\/b> (1886-1982) e <b>Konrad Lorenz<\/b> (1903-1989), cientistas que receberam pr\u00eamio Nobel de Medicina (j\u00e1 que n\u00e3o h\u00e1 pr\u00eamio para etologia) por seus elegant\u00edssimos trabalhos em comportamento animal. Trabalhando predominantemente no campo e por vezes com uma abordagem experimental, estes pesquisadores s\u00e3o considerados os pais da etologia.<\/p>\t\t\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t<div class=\"elementor-element elementor-element-206afea elementor-widget elementor-widget-heading\" data-id=\"206afea\" data-element_type=\"widget\" data-widget_type=\"heading.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t<h2 class=\"elementor-heading-title elementor-size-default\"><span style=\"font-size: 16px;font-weight: 400\">Tinbergen estabeleceu as famosas \u201c<\/span><span style=\"font-size: 16px\"><b>quatro quest\u00f5es so<\/b><\/span><span style=\"font-size: 16px\"><b>bre o comportame<\/b><\/span><span style=\"font-size: 16px\"><b>nto<\/b><\/span><span style=\"font-size: 16px;font-weight: 400\">\u201d:<\/span><\/h2>\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/section>\n\t\t\t\t<section class=\"elementor-section elementor-top-section elementor-element elementor-element-433176a elementor-section-boxed elementor-section-height-default elementor-section-height-default\" data-id=\"433176a\" data-element_type=\"section\">\n\t\t\t\t\t\t<div class=\"elementor-container elementor-column-gap-default\">\n\t\t\t\t\t<div class=\"elementor-column elementor-col-25 elementor-top-column elementor-element elementor-element-9ea2be3\" data-id=\"9ea2be3\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-e83586f elementor-widget elementor-widget-heading\" data-id=\"e83586f\" data-element_type=\"widget\" data-widget_type=\"heading.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t<h2 class=\"elementor-heading-title elementor-size-default\"><h2 class=\"elementor-heading-title elementor-size-default elementor-inline-editing pen\" data-elementor-setting-key=\"title\" data-pen-placeholder=\"Digite aqui...\" style=\"font-size: var( --e-global-typography-primary-font-size );font-family: var( --e-global-typography-primary-font-family ), Sans-serif\"><span style=\"font-weight: 400;font-size: 16px\">Quais s\u00e3o os fatores ex\u00f3genos e end\u00f3genos que desencadeiam um comportamento<\/span><\/h2><\/h2>\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t<div class=\"elementor-column elementor-col-25 elementor-top-column elementor-element elementor-element-b75b214\" data-id=\"b75b214\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-e7cb153 elementor-widget elementor-widget-heading\" data-id=\"e7cb153\" data-element_type=\"widget\" data-widget_type=\"heading.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t<h2 class=\"elementor-heading-title elementor-size-default\"><h2 class=\"elementor-heading-title elementor-size-default elementor-inline-editing pen\" data-elementor-setting-key=\"title\" data-pen-placeholder=\"Digite aqui...\" style=\"font-size: var( --e-global-typography-primary-font-size );font-family: var( --e-global-typography-primary-font-family ), Sans-serif\"><span style=\"font-weight: 400;font-size: 16px\">Como o comportamento se desenvolve em um indiv\u00edduo<\/span><\/h2><\/h2>\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t<div class=\"elementor-column elementor-col-25 elementor-top-column elementor-element elementor-element-764caaf\" data-id=\"764caaf\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-98d3809 elementor-widget elementor-widget-heading\" data-id=\"98d3809\" data-element_type=\"widget\" data-widget_type=\"heading.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t<h2 class=\"elementor-heading-title elementor-size-default\"><h2 class=\"elementor-heading-title elementor-size-default elementor-inline-editing pen\" data-elementor-setting-key=\"title\" data-pen-placeholder=\"Digite aqui...\" style=\"font-size: var( --e-global-typography-primary-font-size );font-family: var( --e-global-typography-primary-font-family ), Sans-serif\"><span style=\"font-weight: 400;font-size: 16px\">Qual o valor adaptativo dos comportamentos<\/span><\/h2><\/h2>\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t<div class=\"elementor-column elementor-col-25 elementor-top-column elementor-element elementor-element-00f3353\" data-id=\"00f3353\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-c1949df elementor-widget elementor-widget-heading\" data-id=\"c1949df\" data-element_type=\"widget\" data-widget_type=\"heading.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t<h2 class=\"elementor-heading-title elementor-size-default\"><h2 class=\"elementor-heading-title elementor-size-default elementor-inline-editing pen\" data-elementor-setting-key=\"title\" data-pen-placeholder=\"Digite aqui...\" style=\"font-size: var( --e-global-typography-primary-font-size );font-family: var( --e-global-typography-primary-font-family ), Sans-serif\"><span style=\"font-weight: 400;font-size: 16px\">Como esses comportamentos se originaram e se modificaram ao longo da hist\u00f3ria evolutiva de uma linhagem.&nbsp;<\/span><\/h2><\/h2>\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/section>\n\t\t\t\t<section class=\"elementor-section elementor-top-section elementor-element elementor-element-1e6e360 elementor-section-boxed elementor-section-height-default elementor-section-height-default\" data-id=\"1e6e360\" data-element_type=\"section\">\n\t\t\t\t\t\t<div class=\"elementor-container elementor-column-gap-default\">\n\t\t\t\t\t<div class=\"elementor-column elementor-col-100 elementor-top-column elementor-element elementor-element-eb9c4dd\" data-id=\"eb9c4dd\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-6a9cb16 elementor-widget elementor-widget-spacer\" data-id=\"6a9cb16\" data-element_type=\"widget\" data-widget_type=\"spacer.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t<div class=\"elementor-spacer\">\n\t\t\t<div class=\"elementor-spacer-inner\"><\/div>\n\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t<div class=\"elementor-element elementor-element-ab2f3b7 elementor-widget elementor-widget-text-editor\" data-id=\"ab2f3b7\" data-element_type=\"widget\" data-widget_type=\"text-editor.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t\t\t<p>\u00a0 \u00a0No sistema de\u00a0<span style=\"font-weight: bold;\">Edward Wilson<\/span>, as duas primeiras quest\u00f5es se encaixariam no que ele chamou de quest\u00f5es proximais e as duas \u00faltimas no que chamamos de quest\u00f5es distais do comportamento, ou ainda as quest\u00f5es do tipo \u201ccomo\u201d e \u201cpor que\u201d de\u00a0<span style=\"font-weight: bold;\">John Alcock<\/span>. Talvez por conta da discuss\u00e3o entre a import\u00e2ncia dos instintos versus a aprendizagem no estabelecimento do comportamento (uma discuss\u00e3o sobre quest\u00f5es proximais), talvez pela falta de compreens\u00e3o dos et\u00f3logos da \u00e9poca sobre a sele\u00e7\u00e3o em n\u00edvel de indiv\u00edduo em oposi\u00e7\u00e3o \u00e0 sele\u00e7\u00e3o de grupo (uma discuss\u00e3o sobre quest\u00f5es distais), quest\u00f5es proximais foram mais estudadas do que quest\u00f5es distais da d\u00e9cada de 50 at\u00e9 in\u00edcio da d\u00e9cada de 70. Ap\u00f3s a aceita\u00e7\u00e3o dos trabalhos de\u00a0<span style=\"font-weight: bold;\">George Williams<\/span>\u00a0(1926-2010) sobre sele\u00e7\u00e3o em n\u00edvel do indiv\u00edduo, o fim da controv\u00e9rsia instinto versus aprendizagem e talvez tamb\u00e9m por influ\u00eancia dos trabalhos de\u00a0<span style=\"font-weight: bold;\">Edward Wilson\u00a0<\/span>(como \u201cSociobiology, 1975\u201d), a quantidade de trabalhos sobre causas distais do comportamento aumentou.<\/p><p>\u00a0 \u00a0Felizmente, nos \u00faltimos anos, ec\u00f3logos comportamentais (que estudam principalmente causas distais), t\u00eam se interessado nas causas proximais do comportamento e neurobi\u00f3logos e psic\u00f3logos cognitivistas (que estudam primariamente as causas proximais) t\u00eam utilizado o m\u00e9todo comparativo e a abordagem evolutiva. A vis\u00e3o de que uma abordagem \u00e9 melhor ou mais importante do que a outra como acontecia nos tempos dos embates cient\u00edficos entre naturalistas e psic\u00f3logos experimentais est\u00e1 obsoleta e n\u00e3o faz mais sentido<\/p>\t\t\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t<div class=\"elementor-element elementor-element-e5e2a54 elementor-widget elementor-widget-spacer\" data-id=\"e5e2a54\" data-element_type=\"widget\" data-widget_type=\"spacer.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t<div class=\"elementor-spacer\">\n\t\t\t<div class=\"elementor-spacer-inner\"><\/div>\n\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/section>\n\t\t\t\t<section class=\"elementor-section elementor-top-section elementor-element elementor-element-71e3055 elementor-section-boxed elementor-section-height-default elementor-section-height-default\" data-id=\"71e3055\" data-element_type=\"section\">\n\t\t\t\t\t\t<div class=\"elementor-container elementor-column-gap-default\">\n\t\t\t\t\t<div class=\"elementor-column elementor-col-100 elementor-top-column elementor-element elementor-element-26128ac\" data-id=\"26128ac\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-1d0fe06 elementor-widget elementor-widget-elementskit-heading\" data-id=\"1d0fe06\" data-element_type=\"widget\" data-widget_type=\"elementskit-heading.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t<div class=\"ekit-wid-con\" ><div class=\"ekit-heading elementskit-section-title-wraper text_left   ekit_heading_tablet-   ekit_heading_mobile-\"><h2 class=\"ekit-heading--title elementskit-section-title \">Como o  <span><span>LESCA<\/span><\/span> se insere nesse contexto?<\/h2><div class=\"ekit_heading_separetor_wraper ekit_heading_elementskit-border-divider\"><div class=\"elementskit-border-divider\"><\/div><\/div><\/div><\/div>\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t<div class=\"elementor-element elementor-element-c454e6b elementor-widget elementor-widget-spacer\" data-id=\"c454e6b\" data-element_type=\"widget\" data-widget_type=\"spacer.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t<div class=\"elementor-spacer\">\n\t\t\t<div class=\"elementor-spacer-inner\"><\/div>\n\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/section>\n\t\t\t\t<section class=\"elementor-section elementor-top-section elementor-element elementor-element-92a7635 elementor-section-boxed elementor-section-height-default elementor-section-height-default\" data-id=\"92a7635\" data-element_type=\"section\">\n\t\t\t\t\t\t<div class=\"elementor-container elementor-column-gap-default\">\n\t\t\t\t\t<div class=\"elementor-column elementor-col-100 elementor-top-column elementor-element elementor-element-4d928bc\" data-id=\"4d928bc\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-2abf11d elementor-widget elementor-widget-text-editor\" data-id=\"2abf11d\" data-element_type=\"widget\" data-widget_type=\"text-editor.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t\t\t<p>\u00a0 \u00a0Em nosso laborat\u00f3rio, investigamos quest\u00f5es essencialmente proximais quando testamos, por exemplo, as modalidades sensoriais importantes para um animal encontrar um parceiro sexual; ou quando trabalhamos com os mecanismos de captura de presas. Estudamos quest\u00f5es evolutivas quando tentamos entender como se deu a evolu\u00e7\u00e3o de estruturas sensoriais ou glandulares em determinadas linhagens; ou ao demostrarmos o valor adaptativo de certos mecanismos defensivos.<\/p><p>\u00a0 \u00a0Metodologicamente, utilizamos t\u00e9cnicas que de certa forma nos aproximam dos naturalistas como observa\u00e7\u00f5es e descri\u00e7\u00f5es do comportamento no campo e t\u00e9cnicas mais parecidas com as da psicologia experimental quando realizamos experimentos em condi\u00e7\u00f5es rigorosamente controladas de laborat\u00f3rio. Adicionalmente, realizamos estudos morfol\u00f3gicos e qu\u00edmicos que nos auxiliam a responder nossas perguntas. Rejeitamos a id\u00e9ia de que determinada abordagem ou t\u00e9cnica \u00e9 mais importante do que outra. Pensamos que h\u00e1 problemas biologicamente interessantes para serem resolvidos tanto nas causas proximais quanto distais do comportamento e que m\u00e9todos experimentais ou descritivos s\u00e3o igualmente \u00fateis dependendo de qual pergunta deve ser respondida. A uni\u00e3o entre estas abordagens e t\u00e9cnicas s\u00f3 tem a contribuir a uma melhor compreens\u00e3o dos mais diversos aspectos desses animais que tanto nos fascinam.<\/p>\t\t\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/section>\n\t\t\t\t<section class=\"elementor-section elementor-top-section elementor-element elementor-element-1b9a07a elementor-section-boxed elementor-section-height-default elementor-section-height-default\" data-id=\"1b9a07a\" data-element_type=\"section\">\n\t\t\t\t\t\t<div class=\"elementor-container elementor-column-gap-default\">\n\t\t\t\t\t<div class=\"elementor-column elementor-col-100 elementor-top-column elementor-element elementor-element-3d37adf\" data-id=\"3d37adf\" data-element_type=\"column\">\n\t\t\t<div class=\"elementor-widget-wrap elementor-element-populated\">\n\t\t\t\t\t\t<div class=\"elementor-element elementor-element-7410043 elementor-widget elementor-widget-accordion\" data-id=\"7410043\" data-element_type=\"widget\" data-widget_type=\"accordion.default\">\n\t\t\t\t<div class=\"elementor-widget-container\">\n\t\t\t\t\t<div class=\"elementor-accordion\">\n\t\t\t\t\t\t\t<div class=\"elementor-accordion-item\">\n\t\t\t\t\t<div id=\"elementor-tab-title-1211\" class=\"elementor-tab-title\" data-tab=\"1\" role=\"button\" aria-controls=\"elementor-tab-content-1211\" aria-expanded=\"false\">\n\t\t\t\t\t\t\t\t\t\t\t\t\t<span class=\"elementor-accordion-icon elementor-accordion-icon-left\" aria-hidden=\"true\">\n\t\t\t\t\t\t\t\t\t\t\t\t\t\t\t<span class=\"elementor-accordion-icon-closed\"><i class=\"fas fa-plus\"><\/i><\/span>\n\t\t\t\t\t\t\t\t<span class=\"elementor-accordion-icon-opened\"><i class=\"fas fa-minus\"><\/i><\/span>\n\t\t\t\t\t\t\t\t\t\t\t\t\t\t<\/span>\n\t\t\t\t\t\t\t\t\t\t\t\t<a class=\"elementor-accordion-title\" tabindex=\"0\">Refer\u00eancias<\/a>\n\t\t\t\t\t<\/div>\n\t\t\t\t\t<div id=\"elementor-tab-content-1211\" class=\"elementor-tab-content elementor-clearfix\" data-tab=\"1\" role=\"region\" aria-labelledby=\"elementor-tab-title-1211\"><p><br>Alcock J 2003. A textbook history of animal behavior. Animal Behaviour 65: 3-10.<\/p>\n<p>Alcock J &amp; P Sherman 1994. The Utility of the Proximate-Ultimate Dichotomy in Ethology. Ethology 96: 58-62.<\/p>\n<p>C\u00e9zilly F 2001.Things have(n\u2019t) changed! Behavioural Processes 55: 65-66.<\/p>\n<p>Danchin E, LA Giraldeau &amp; F C\u00e9zilly 2008. Behavioural Ecology: An evolutionary perspective on behaviour. Oxford University Press.<\/p>\n<p>Hogan JA 2005. Causation: the study of behavioural mechanisms. Animal Biology 55: 323-341.<\/p>\n<p>Lehner PN 1996. Handbook of ethological methods, 2nd Ed. Cambridge University Press.<\/p>\n<p>Ryan MJ 2005.The evolution of behaviour, and integrating it towards a complete and correct understanding of behavioural biology. Animal Biology 55: 419-439.<\/p>\n<p>Schoenfeld WN &amp; HS Baron 1965. Ethology and Experimental Psychology. Science 147: 634-635.<\/p>\n<p>Segerstrale U 2000. Defenders of the truth: the battle for science in the sociobiology debate and beyond. Oxford University Press.<\/p>\n<p>Thierry B 2007. Behaviorology divided: Shall we continue? Behaviour 144: 861-878.<\/p>\n<p>Thierry B 2010. Darwin as a student of behavior. Comptes Rendus Biologies 333: 188-196.<\/p>\n<p>Tinbergen N 1963. On aims and methods of ethology. Zeitschrift f\u00fcr Tierpsychologie 20: 410-433.<\/p>\n<p>Vauclair J 1996. Animal cognition: an introduction to modern comparative psychology. Harvard University Press.<\/p>\n<p>Williams GC 1966. Adaptation and Natural Selection. Princeton, New Jersey: Princeton University Press.<\/p>\n<p>Wilson EO 1975. Sociobiology: the New Synthesis. Cambridge, Massachusetts: Harvard University Press<\/p><\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/div>\n\t\t\t\t\t<\/div>\n\t\t<\/section>\n\t\t\t\t<\/div>\n\t\t","protected":false},"excerpt":{"rendered":"<p>\u00a0 \u00a0Estudo do comportamento animal e o LESCANeste texto farei um brev\u00edssimo hist\u00f3rico de estudos sobre comportamento animal e ao final tentarei encaixar o LESCA nesse contexto em termos de abordagem e m\u00e9todos. \u00a0 \u00a0Do s\u00e9culo XVIII at\u00e9 o s\u00e9culo XIX, duas escolas de pensamento discutiram avidamente sobre qual a maneira mais adequada de se &hellip; <\/p>\n<p class=\"link-more\"><a href=\"https:\/\/sites.usp.br\/lesca\/comportamento-animal\/\" class=\"more-link\">Continue lendo<span class=\"screen-reader-text\"> &#8220;Comportamento animal&#8221;<\/span><\/a><\/p>\n","protected":false},"author":24477,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"footnotes":"","_links_to":"","_links_to_target":""},"class_list":["post-868","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/sites.usp.br\/lesca\/wp-json\/wp\/v2\/pages\/868","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/sites.usp.br\/lesca\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/sites.usp.br\/lesca\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/sites.usp.br\/lesca\/wp-json\/wp\/v2\/users\/24477"}],"replies":[{"embeddable":true,"href":"https:\/\/sites.usp.br\/lesca\/wp-json\/wp\/v2\/comments?post=868"}],"version-history":[{"count":4,"href":"https:\/\/sites.usp.br\/lesca\/wp-json\/wp\/v2\/pages\/868\/revisions"}],"predecessor-version":[{"id":1586,"href":"https:\/\/sites.usp.br\/lesca\/wp-json\/wp\/v2\/pages\/868\/revisions\/1586"}],"wp:attachment":[{"href":"https:\/\/sites.usp.br\/lesca\/wp-json\/wp\/v2\/media?parent=868"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}